Functional redundancy dampens precipitation change impacts on species-rich invertebrate communities across the Neotropics

Autores: Céréghino, Régis; Trzcinski, Mark Kurtis; MacDonald, A. Andrew M.; Marino, Nicholas A. C.; Acosta Mercado, Dimaris; Leroy, Céline; Corbara, Bruno; Romero, Gustavo Q.; Farjalla, Vinicius F.; Barberis, Ignacio Martín; Dézerald, Olivier; Hammill, Edd; Atwood, Trisha B.; Piccoli, Gustavo C. O.; Ospina Bautista, Fabiola; Carrias, Jean François; Leal, Juliana S.; Montero, Guillermo Alberto; Antiqueira, Pablo A. P.; Freire, Rodrigo Manuel; Realpe, Emilio; Amundrud, Sarah L.; de Omena, Paula M.; Campos, Alice B. A.; Srivastava, Diane S.
Año de publicación: 2022
Idioma: inglés
Tipo de recurso: artículo
Estado: versión publicada
Descripción: Animal community responses to extreme climate events can be predicted from the functional traits represented within communities. However, it is unclear whether geographic variation in the response of functional community structure to climate change is primarily driven by physiological matching to local conditions (local adaptation hypothesis) or by differences between species pools in functional redundancy (insurance hypothesis). We conducted a coordinated experiment to understand how aquatic invertebrate traits mediate the responses of multitrophic communities to changes in the quantity and evenness of rainfall in 180 natural freshwater microcosms (tank bromeliads) distributed across six sites from 18°N in the Caribbean to 29°S in South America. At each site, we manipulated the mean and dispersion of the daily amount of rainfall that entered tank bromeliads over a 2-month period. Manipulations covered a response surface representing 50% to 200% of the dispersion of daily rainfall crossed with 10% to 300% of the mean amounts of rainfall. The response of functional community structure to precipitation regimes differed across sites. These geographic differences were not consistent with the local adaptation hypothesis, as responses did not correlate with the current amplitude in precipitation. Geographic differences in community responses were consistent with the insurance hypothesis: sites with the lowest functional redundancy in their species pools had the strongest response to a gradient in hydrological variability induced by uneven precipitation. In such sites, an increase in the hydrologic variability induced a shift from communities with both pelagic and benthic traits using both green and brown energy channels to strictly benthic, brown energy communities. Our results predict uneven impacts of precipitation change on community structure and energy channels within communities across Neotropical regions. This geographic variation is due more to differences in the size and redundancy of species pools than to local adaptation. Strategies for climate change adaptation should thus seek to identify and preserve functionally unique species and their habitats. Read the free Plain Language Summary for this article on the Journal blog.
Fil: Céréghino, Régis. Universite de Toulouse; Francia
Fil: Trzcinski, Mark Kurtis. University of British Columbia; Canadá
Fil: MacDonald, A. Andrew M.. Centre For Biodiversity Science; Canadá
Fil: Marino, Nicholas A. C.. Universidade Federal do Rio de Janeiro; Brasil
Fil: Acosta Mercado, Dimaris. Universidad de Puerto Rico; Puerto Rico
Fil: Leroy, Céline. Université Montpellier II; Francia
Fil: Corbara, Bruno. No especifíca;
Fil: Romero, Gustavo Q.. Universidade Estadual de Campinas; Brasil
Fil: Farjalla, Vinicius F.. Universidade Federal do Rio de Janeiro; Brasil
Fil: Barberis, Ignacio Martín. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico Conicet - Rosario. Instituto de Investigaciones en Ciencias Agrarias de Rosario. Universidad Nacional de Rosario. Facultad de Ciencias Agrarias. Instituto de Investigaciones en Ciencias Agrarias de Rosario; Argentina
Fil: Dézerald, Olivier. Universite de Toulouse; Francia
Fil: Hammill, Edd. University of Utah; Estados Unidos
Fil: Atwood, Trisha B.. University of Utah; Estados Unidos
Fil: Piccoli, Gustavo C. O.. Universidade de Sao Paulo; Brasil
Fil: Ospina Bautista, Fabiola. Universidad de Caldas; Colombia
Fil: Carrias, Jean François. No especifíca;
Fil: Leal, Juliana S.. Universidade Federal do Rio de Janeiro; Brasil
Fil: Montero, Guillermo Alberto. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico Conicet - Rosario. Instituto de Investigaciones en Ciencias Agrarias de Rosario. Universidad Nacional de Rosario. Facultad de Ciencias Agrarias. Instituto de Investigaciones en Ciencias Agrarias de Rosario; Argentina
Fil: Antiqueira, Pablo A. P.. Universidade Estadual de Campinas; Brasil
Fil: Freire, Rodrigo Manuel. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico Conicet - Rosario. Instituto de Investigaciones en Ciencias Agrarias de Rosario. Universidad Nacional de Rosario. Facultad de Ciencias Agrarias. Instituto de Investigaciones en Ciencias Agrarias de Rosario; Argentina
Fil: Realpe, Emilio. Universidad de Caldas; Colombia
Fil: Amundrud, Sarah L.. Biodiversity Research Centre; Canadá
Fil: de Omena, Paula M.. Universidade Estadual de Campinas; Brasil
Fil: Campos, Alice B. A.. Universidade Federal do Rio de Janeiro; Brasil
Fil: Srivastava, Diane S.. Biodiversity Research Centre; Canadá
Materia: FRESHWATER
FUNCTIONAL TRAITS
HYDROLOGY
INSURANCE HYPOTHESIS
PRECIPITATION
SPECIES RICHNESS
Nivel de accesibilidad: acceso abierto
Condiciones de uso: https://creativecommons.org/licenses/by/2.5/ar/
Repositorio
Institución: Consejo Nacional de Investigaciones Científicas y Técnicas
OAI Identificador: oai:ri.conicet.gov.ar:11336/213390

Acceder

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spelling	Functional redundancy dampens precipitation change impacts on species-rich invertebrate communities across the NeotropicsCéréghino, RégisTrzcinski, Mark KurtisMacDonald, A. Andrew M.Marino, Nicholas A. C.Acosta Mercado, DimarisLeroy, CélineCorbara, BrunoRomero, Gustavo Q.Farjalla, Vinicius F.Barberis, Ignacio MartínDézerald, OlivierHammill, EddAtwood, Trisha B.Piccoli, Gustavo C. O.Ospina Bautista, FabiolaCarrias, Jean FrançoisLeal, Juliana S.Montero, Guillermo AlbertoAntiqueira, Pablo A. P.Freire, Rodrigo ManuelRealpe, EmilioAmundrud, Sarah L.de Omena, Paula M.Campos, Alice B. A.Srivastava, Diane S.FRESHWATERFUNCTIONAL TRAITSHYDROLOGYINSURANCE HYPOTHESISPRECIPITATIONSPECIES RICHNESShttps://purl.org/becyt/ford/1.6https://purl.org/becyt/ford/1Animal community responses to extreme climate events can be predicted from the functional traits represented within communities. However, it is unclear whether geographic variation in the response of functional community structure to climate change is primarily driven by physiological matching to local conditions (local adaptation hypothesis) or by differences between species pools in functional redundancy (insurance hypothesis). We conducted a coordinated experiment to understand how aquatic invertebrate traits mediate the responses of multitrophic communities to changes in the quantity and evenness of rainfall in 180 natural freshwater microcosms (tank bromeliads) distributed across six sites from 18°N in the Caribbean to 29°S in South America. At each site, we manipulated the mean and dispersion of the daily amount of rainfall that entered tank bromeliads over a 2-month period. Manipulations covered a response surface representing 50% to 200% of the dispersion of daily rainfall crossed with 10% to 300% of the mean amounts of rainfall. The response of functional community structure to precipitation regimes differed across sites. These geographic differences were not consistent with the local adaptation hypothesis, as responses did not correlate with the current amplitude in precipitation. Geographic differences in community responses were consistent with the insurance hypothesis: sites with the lowest functional redundancy in their species pools had the strongest response to a gradient in hydrological variability induced by uneven precipitation. In such sites, an increase in the hydrologic variability induced a shift from communities with both pelagic and benthic traits using both green and brown energy channels to strictly benthic, brown energy communities. Our results predict uneven impacts of precipitation change on community structure and energy channels within communities across Neotropical regions. This geographic variation is due more to differences in the size and redundancy of species pools than to local adaptation. Strategies for climate change adaptation should thus seek to identify and preserve functionally unique species and their habitats. Read the free Plain Language Summary for this article on the Journal blog.Fil: Céréghino, Régis. Universite de Toulouse; FranciaFil: Trzcinski, Mark Kurtis. University of British Columbia; CanadáFil: MacDonald, A. Andrew M.. Centre For Biodiversity Science; CanadáFil: Marino, Nicholas A. C.. Universidade Federal do Rio de Janeiro; BrasilFil: Acosta Mercado, Dimaris. Universidad de Puerto Rico; Puerto RicoFil: Leroy, Céline. Université Montpellier II; FranciaFil: Corbara, Bruno. No especifíca;Fil: Romero, Gustavo Q.. Universidade Estadual de Campinas; BrasilFil: Farjalla, Vinicius F.. Universidade Federal do Rio de Janeiro; BrasilFil: Barberis, Ignacio Martín. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico Conicet - Rosario. Instituto de Investigaciones en Ciencias Agrarias de Rosario. Universidad Nacional de Rosario. Facultad de Ciencias Agrarias. Instituto de Investigaciones en Ciencias Agrarias de Rosario; ArgentinaFil: Dézerald, Olivier. Universite de Toulouse; FranciaFil: Hammill, Edd. University of Utah; Estados UnidosFil: Atwood, Trisha B.. University of Utah; Estados UnidosFil: Piccoli, Gustavo C. O.. Universidade de Sao Paulo; BrasilFil: Ospina Bautista, Fabiola. Universidad de Caldas; ColombiaFil: Carrias, Jean François. No especifíca;Fil: Leal, Juliana S.. Universidade Federal do Rio de Janeiro; BrasilFil: Montero, Guillermo Alberto. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico Conicet - Rosario. Instituto de Investigaciones en Ciencias Agrarias de Rosario. Universidad Nacional de Rosario. Facultad de Ciencias Agrarias. Instituto de Investigaciones en Ciencias Agrarias de Rosario; ArgentinaFil: Antiqueira, Pablo A. P.. Universidade Estadual de Campinas; BrasilFil: Freire, Rodrigo Manuel. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico Conicet - Rosario. Instituto de Investigaciones en Ciencias Agrarias de Rosario. Universidad Nacional de Rosario. Facultad de Ciencias Agrarias. Instituto de Investigaciones en Ciencias Agrarias de Rosario; ArgentinaFil: Realpe, Emilio. Universidad de Caldas; ColombiaFil: Amundrud, Sarah L.. Biodiversity Research Centre; CanadáFil: de Omena, Paula M.. Universidade Estadual de Campinas; BrasilFil: Campos, Alice B. A.. Universidade Federal do Rio de Janeiro; BrasilFil: Srivastava, Diane S.. Biodiversity Research Centre; CanadáWiley Blackwell Publishing, Inc2022-04info:eu-repo/semantics/articleinfo:eu-repo/semantics/publishedVersionhttp://purl.org/coar/resource_type/c_6501info:ar-repo/semantics/articuloapplication/pdfapplication/pdfapplication/pdfhttp://hdl.handle.net/11336/213390Céréghino, Régis; Trzcinski, Mark Kurtis; MacDonald, A. Andrew M.; Marino, Nicholas A. C.; Acosta Mercado, Dimaris; et al.; Functional redundancy dampens precipitation change impacts on species-rich invertebrate communities across the Neotropics; Wiley Blackwell Publishing, Inc; Functional Ecology; 36; 7; 4-2022; 1559-15720269-8463CONICET DigitalCONICETenginfo:eu-repo/semantics/altIdentifier/url/https://onlinelibrary.wiley.com/doi/10.1111/1365-2435.14048info:eu-repo/semantics/altIdentifier/doi/10.1111/1365-2435.14048info:eu-repo/semantics/openAccesshttps://creativecommons.org/licenses/by/2.5/ar/reponame:CONICET Digital (CONICET)instname:Consejo Nacional de Investigaciones Científicas y Técnicas2025-10-15T14:49:47Zoai:ri.conicet.gov.ar:11336/213390instacron:CONICETInstitucionalhttp://ri.conicet.gov.ar/Organismo científico-tecnológicoNo correspondehttp://ri.conicet.gov.ar/oai/requestdasensio@conicet.gov.ar; lcarlino@conicet.gov.arArgentinaNo correspondeNo correspondeNo correspondeopendoar:34982025-10-15 14:49:48.057CONICET Digital (CONICET) - Consejo Nacional de Investigaciones Científicas y Técnicasfalse
dc.title.none.fl_str_mv	Functional redundancy dampens precipitation change impacts on species-rich invertebrate communities across the Neotropics
title	Functional redundancy dampens precipitation change impacts on species-rich invertebrate communities across the Neotropics
spellingShingle	Functional redundancy dampens precipitation change impacts on species-rich invertebrate communities across the Neotropics Céréghino, Régis FRESHWATER FUNCTIONAL TRAITS HYDROLOGY INSURANCE HYPOTHESIS PRECIPITATION SPECIES RICHNESS
title_short	Functional redundancy dampens precipitation change impacts on species-rich invertebrate communities across the Neotropics
title_full	Functional redundancy dampens precipitation change impacts on species-rich invertebrate communities across the Neotropics
title_fullStr	Functional redundancy dampens precipitation change impacts on species-rich invertebrate communities across the Neotropics
title_full_unstemmed	Functional redundancy dampens precipitation change impacts on species-rich invertebrate communities across the Neotropics
title_sort	Functional redundancy dampens precipitation change impacts on species-rich invertebrate communities across the Neotropics
dc.creator.none.fl_str_mv	Céréghino, Régis Trzcinski, Mark Kurtis MacDonald, A. Andrew M. Marino, Nicholas A. C. Acosta Mercado, Dimaris Leroy, Céline Corbara, Bruno Romero, Gustavo Q. Farjalla, Vinicius F. Barberis, Ignacio Martín Dézerald, Olivier Hammill, Edd Atwood, Trisha B. Piccoli, Gustavo C. O. Ospina Bautista, Fabiola Carrias, Jean François Leal, Juliana S. Montero, Guillermo Alberto Antiqueira, Pablo A. P. Freire, Rodrigo Manuel Realpe, Emilio Amundrud, Sarah L. de Omena, Paula M. Campos, Alice B. A. Srivastava, Diane S.
author	Céréghino, Régis
author_facet	Céréghino, Régis Trzcinski, Mark Kurtis MacDonald, A. Andrew M. Marino, Nicholas A. C. Acosta Mercado, Dimaris Leroy, Céline Corbara, Bruno Romero, Gustavo Q. Farjalla, Vinicius F. Barberis, Ignacio Martín Dézerald, Olivier Hammill, Edd Atwood, Trisha B. Piccoli, Gustavo C. O. Ospina Bautista, Fabiola Carrias, Jean François Leal, Juliana S. Montero, Guillermo Alberto Antiqueira, Pablo A. P. Freire, Rodrigo Manuel Realpe, Emilio Amundrud, Sarah L. de Omena, Paula M. Campos, Alice B. A. Srivastava, Diane S.
author_role	author
author2	Trzcinski, Mark Kurtis MacDonald, A. Andrew M. Marino, Nicholas A. C. Acosta Mercado, Dimaris Leroy, Céline Corbara, Bruno Romero, Gustavo Q. Farjalla, Vinicius F. Barberis, Ignacio Martín Dézerald, Olivier Hammill, Edd Atwood, Trisha B. Piccoli, Gustavo C. O. Ospina Bautista, Fabiola Carrias, Jean François Leal, Juliana S. Montero, Guillermo Alberto Antiqueira, Pablo A. P. Freire, Rodrigo Manuel Realpe, Emilio Amundrud, Sarah L. de Omena, Paula M. Campos, Alice B. A. Srivastava, Diane S.
author2_role	author author author author author author author author author author author author author author author author author author author author author author author author
dc.subject.none.fl_str_mv	FRESHWATER FUNCTIONAL TRAITS HYDROLOGY INSURANCE HYPOTHESIS PRECIPITATION SPECIES RICHNESS
topic	FRESHWATER FUNCTIONAL TRAITS HYDROLOGY INSURANCE HYPOTHESIS PRECIPITATION SPECIES RICHNESS
purl_subject.fl_str_mv	https://purl.org/becyt/ford/1.6 https://purl.org/becyt/ford/1
dc.description.none.fl_txt_mv	Animal community responses to extreme climate events can be predicted from the functional traits represented within communities. However, it is unclear whether geographic variation in the response of functional community structure to climate change is primarily driven by physiological matching to local conditions (local adaptation hypothesis) or by differences between species pools in functional redundancy (insurance hypothesis). We conducted a coordinated experiment to understand how aquatic invertebrate traits mediate the responses of multitrophic communities to changes in the quantity and evenness of rainfall in 180 natural freshwater microcosms (tank bromeliads) distributed across six sites from 18°N in the Caribbean to 29°S in South America. At each site, we manipulated the mean and dispersion of the daily amount of rainfall that entered tank bromeliads over a 2-month period. Manipulations covered a response surface representing 50% to 200% of the dispersion of daily rainfall crossed with 10% to 300% of the mean amounts of rainfall. The response of functional community structure to precipitation regimes differed across sites. These geographic differences were not consistent with the local adaptation hypothesis, as responses did not correlate with the current amplitude in precipitation. Geographic differences in community responses were consistent with the insurance hypothesis: sites with the lowest functional redundancy in their species pools had the strongest response to a gradient in hydrological variability induced by uneven precipitation. In such sites, an increase in the hydrologic variability induced a shift from communities with both pelagic and benthic traits using both green and brown energy channels to strictly benthic, brown energy communities. Our results predict uneven impacts of precipitation change on community structure and energy channels within communities across Neotropical regions. This geographic variation is due more to differences in the size and redundancy of species pools than to local adaptation. Strategies for climate change adaptation should thus seek to identify and preserve functionally unique species and their habitats. Read the free Plain Language Summary for this article on the Journal blog. Fil: Céréghino, Régis. Universite de Toulouse; Francia Fil: Trzcinski, Mark Kurtis. University of British Columbia; Canadá Fil: MacDonald, A. Andrew M.. Centre For Biodiversity Science; Canadá Fil: Marino, Nicholas A. C.. Universidade Federal do Rio de Janeiro; Brasil Fil: Acosta Mercado, Dimaris. Universidad de Puerto Rico; Puerto Rico Fil: Leroy, Céline. Université Montpellier II; Francia Fil: Corbara, Bruno. No especifíca; Fil: Romero, Gustavo Q.. Universidade Estadual de Campinas; Brasil Fil: Farjalla, Vinicius F.. Universidade Federal do Rio de Janeiro; Brasil Fil: Barberis, Ignacio Martín. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico Conicet - Rosario. Instituto de Investigaciones en Ciencias Agrarias de Rosario. Universidad Nacional de Rosario. Facultad de Ciencias Agrarias. Instituto de Investigaciones en Ciencias Agrarias de Rosario; Argentina Fil: Dézerald, Olivier. Universite de Toulouse; Francia Fil: Hammill, Edd. University of Utah; Estados Unidos Fil: Atwood, Trisha B.. University of Utah; Estados Unidos Fil: Piccoli, Gustavo C. O.. Universidade de Sao Paulo; Brasil Fil: Ospina Bautista, Fabiola. Universidad de Caldas; Colombia Fil: Carrias, Jean François. No especifíca; Fil: Leal, Juliana S.. Universidade Federal do Rio de Janeiro; Brasil Fil: Montero, Guillermo Alberto. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico Conicet - Rosario. Instituto de Investigaciones en Ciencias Agrarias de Rosario. Universidad Nacional de Rosario. Facultad de Ciencias Agrarias. Instituto de Investigaciones en Ciencias Agrarias de Rosario; Argentina Fil: Antiqueira, Pablo A. P.. Universidade Estadual de Campinas; Brasil Fil: Freire, Rodrigo Manuel. Consejo Nacional de Investigaciones Científicas y Técnicas. Centro Científico Tecnológico Conicet - Rosario. Instituto de Investigaciones en Ciencias Agrarias de Rosario. Universidad Nacional de Rosario. Facultad de Ciencias Agrarias. Instituto de Investigaciones en Ciencias Agrarias de Rosario; Argentina Fil: Realpe, Emilio. Universidad de Caldas; Colombia Fil: Amundrud, Sarah L.. Biodiversity Research Centre; Canadá Fil: de Omena, Paula M.. Universidade Estadual de Campinas; Brasil Fil: Campos, Alice B. A.. Universidade Federal do Rio de Janeiro; Brasil Fil: Srivastava, Diane S.. Biodiversity Research Centre; Canadá
description	Animal community responses to extreme climate events can be predicted from the functional traits represented within communities. However, it is unclear whether geographic variation in the response of functional community structure to climate change is primarily driven by physiological matching to local conditions (local adaptation hypothesis) or by differences between species pools in functional redundancy (insurance hypothesis). We conducted a coordinated experiment to understand how aquatic invertebrate traits mediate the responses of multitrophic communities to changes in the quantity and evenness of rainfall in 180 natural freshwater microcosms (tank bromeliads) distributed across six sites from 18°N in the Caribbean to 29°S in South America. At each site, we manipulated the mean and dispersion of the daily amount of rainfall that entered tank bromeliads over a 2-month period. Manipulations covered a response surface representing 50% to 200% of the dispersion of daily rainfall crossed with 10% to 300% of the mean amounts of rainfall. The response of functional community structure to precipitation regimes differed across sites. These geographic differences were not consistent with the local adaptation hypothesis, as responses did not correlate with the current amplitude in precipitation. Geographic differences in community responses were consistent with the insurance hypothesis: sites with the lowest functional redundancy in their species pools had the strongest response to a gradient in hydrological variability induced by uneven precipitation. In such sites, an increase in the hydrologic variability induced a shift from communities with both pelagic and benthic traits using both green and brown energy channels to strictly benthic, brown energy communities. Our results predict uneven impacts of precipitation change on community structure and energy channels within communities across Neotropical regions. This geographic variation is due more to differences in the size and redundancy of species pools than to local adaptation. Strategies for climate change adaptation should thus seek to identify and preserve functionally unique species and their habitats. Read the free Plain Language Summary for this article on the Journal blog.
publishDate	2022
dc.date.none.fl_str_mv	2022-04
dc.type.none.fl_str_mv	info:eu-repo/semantics/article info:eu-repo/semantics/publishedVersion http://purl.org/coar/resource_type/c_6501 info:ar-repo/semantics/articulo
format	article
status_str	publishedVersion
dc.identifier.none.fl_str_mv	http://hdl.handle.net/11336/213390 Céréghino, Régis; Trzcinski, Mark Kurtis; MacDonald, A. Andrew M.; Marino, Nicholas A. C.; Acosta Mercado, Dimaris; et al.; Functional redundancy dampens precipitation change impacts on species-rich invertebrate communities across the Neotropics; Wiley Blackwell Publishing, Inc; Functional Ecology; 36; 7; 4-2022; 1559-1572 0269-8463 CONICET Digital CONICET
url	http://hdl.handle.net/11336/213390
identifier_str_mv	Céréghino, Régis; Trzcinski, Mark Kurtis; MacDonald, A. Andrew M.; Marino, Nicholas A. C.; Acosta Mercado, Dimaris; et al.; Functional redundancy dampens precipitation change impacts on species-rich invertebrate communities across the Neotropics; Wiley Blackwell Publishing, Inc; Functional Ecology; 36; 7; 4-2022; 1559-1572 0269-8463 CONICET Digital CONICET
dc.language.none.fl_str_mv	eng
language	eng
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dc.publisher.none.fl_str_mv	Wiley Blackwell Publishing, Inc
publisher.none.fl_str_mv	Wiley Blackwell Publishing, Inc
dc.source.none.fl_str_mv	reponame:CONICET Digital (CONICET) instname:Consejo Nacional de Investigaciones Científicas y Técnicas
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repository.mail.fl_str_mv	dasensio@conicet.gov.ar; lcarlino@conicet.gov.ar
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Functional redundancy dampens precipitation change impacts on species-rich invertebrate communities across the Neotropics

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