Testudinid turtle remains from the Late Miocene palaeo-island of Gargano, Italy, and an overview of Mediterranean insular tortoises

Autores
Georgalis, Georgios L.; Vlachos, Evangelos; Camanni, Francesca; Villa, Andrea; Pavia, Marco; van den Hoek Ostende, Lars W.; Delfino, Massimo
Año de publicación
2025
Idioma
inglés
Tipo de recurso
artículo
Estado
versión publicada
Descripción
The Late Miocene Gargano “Terre Rosse” in southern Italy has yielded a diverse vertebrate fauna that comprises an array of endemic taxa, including forms characterized by extreme sizes and unique morphologies. We here document new testudinid fossil remains from this insular assemblage. The material comprises a mandible, abundant (though mostly fragmentary) shell remains, appendicular elements, and a few vertebrae, originating from different localities within the Gargano assemblage. Based on its femur morphology, we tentatively refer the Gargano form to Solitudo, a late Neogene and Quaternary insular genus, otherwise known from a few Mediterranean islands. Unfortunately, the incomplete preservation of the trochanters hinders observation of the main diagnostic character of Solitudo, however, the similarities in the femoral head and diaphysis allow an open identification of the Gargano material as cf. Solitudo sp. Still, the Gargano tortoise offers new anatomical information on the Mediterranean insular tortoises. An interesting feature of the few preserved plastral elements is the presence of sulci as distinct raised ridges, a character that has been reported in other insular extant and extinct tortoises as well. The preserved hyoplastron has a medially straight and laterally concave humeropectoral sulcus and a wide axillary scute, pointing to some resemblance to geochelonans. Moreover, the morphology of the mandible from Gargano, which has its lingual serration reaching towards the symphyseal area, is also indicative of geochelonan affinities. If our identification is correct, the Gargano form would correspond to the earliest occurrence of Solitudo, denoting that this insular testudinid already radiated during the Late Miocene. A thorough overview of the diversity and taxonomy of insular testudinids from the Mediterranean islands is provided.
Fil: Georgalis, Georgios L.. Polish Academy Of Sciences. Institute Of Paleobiology; Argentina
Fil: Vlachos, Evangelos. Museo Paleontológico Egidio Feruglio; Argentina. Consejo Nacional de Investigaciones Científicas y Técnicas; Argentina
Fil: Camanni, Francesca. Università di Torino; Italia
Fil: Villa, Andrea. Institut Català de Paleontologia Miquel Crusafont.; España
Fil: Pavia, Marco. Università di Torino; Italia
Fil: van den Hoek Ostende, Lars W.. Naturalis Biodiversity Center; Países Bajos
Fil: Delfino, Massimo. Università di Torino; Italia
Materia
TURTLES
TESTUDINIDAE
ANATOMY
TAXONOMY
PALAEOGEOGRAPHY
INSULAR FAUNAS
Nivel de accesibilidad
acceso abierto
Condiciones de uso
https://creativecommons.org/licenses/by/2.5/ar/
Repositorio
CONICET Digital (CONICET)
Institución
Consejo Nacional de Investigaciones Científicas y Técnicas
OAI Identificador
oai:ri.conicet.gov.ar:11336/274803
Acceder
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network_name_str CONICET Digital (CONICET)
spelling Testudinid turtle remains from the Late Miocene palaeo-island of Gargano, Italy, and an overview of Mediterranean insular tortoisesGeorgalis, Georgios L.Vlachos, EvangelosCamanni, FrancescaVilla, AndreaPavia, Marcovan den Hoek Ostende, Lars W.Delfino, MassimoTURTLESTESTUDINIDAEANATOMYTAXONOMYPALAEOGEOGRAPHYINSULAR FAUNAShttps://purl.org/becyt/ford/1.5https://purl.org/becyt/ford/1The Late Miocene Gargano “Terre Rosse” in southern Italy has yielded a diverse vertebrate fauna that comprises an array of endemic taxa, including forms characterized by extreme sizes and unique morphologies. We here document new testudinid fossil remains from this insular assemblage. The material comprises a mandible, abundant (though mostly fragmentary) shell remains, appendicular elements, and a few vertebrae, originating from different localities within the Gargano assemblage. Based on its femur morphology, we tentatively refer the Gargano form to Solitudo, a late Neogene and Quaternary insular genus, otherwise known from a few Mediterranean islands. Unfortunately, the incomplete preservation of the trochanters hinders observation of the main diagnostic character of Solitudo, however, the similarities in the femoral head and diaphysis allow an open identification of the Gargano material as cf. Solitudo sp. Still, the Gargano tortoise offers new anatomical information on the Mediterranean insular tortoises. An interesting feature of the few preserved plastral elements is the presence of sulci as distinct raised ridges, a character that has been reported in other insular extant and extinct tortoises as well. The preserved hyoplastron has a medially straight and laterally concave humeropectoral sulcus and a wide axillary scute, pointing to some resemblance to geochelonans. Moreover, the morphology of the mandible from Gargano, which has its lingual serration reaching towards the symphyseal area, is also indicative of geochelonan affinities. If our identification is correct, the Gargano form would correspond to the earliest occurrence of Solitudo, denoting that this insular testudinid already radiated during the Late Miocene. A thorough overview of the diversity and taxonomy of insular testudinids from the Mediterranean islands is provided.Fil: Georgalis, Georgios L.. Polish Academy Of Sciences. Institute Of Paleobiology; ArgentinaFil: Vlachos, Evangelos. Museo Paleontológico Egidio Feruglio; Argentina. Consejo Nacional de Investigaciones Científicas y Técnicas; ArgentinaFil: Camanni, Francesca. Università di Torino; ItaliaFil: Villa, Andrea. Institut Català de Paleontologia Miquel Crusafont.; EspañaFil: Pavia, Marco. Università di Torino; ItaliaFil: van den Hoek Ostende, Lars W.. Naturalis Biodiversity Center; Países BajosFil: Delfino, Massimo. Università di Torino; ItaliaSpringer2025-09info:eu-repo/semantics/articleinfo:eu-repo/semantics/publishedVersionhttp://purl.org/coar/resource_type/c_6501info:ar-repo/semantics/articuloapplication/pdfapplication/pdfhttp://hdl.handle.net/11336/274803Georgalis, Georgios L.; Vlachos, Evangelos; Camanni, Francesca; Villa, Andrea; Pavia, Marco; et al.; Testudinid turtle remains from the Late Miocene palaeo-island of Gargano, Italy, and an overview of Mediterranean insular tortoises; Springer; Swiss Journal of Palaeontology; 144; 1; 9-2025; 1-241664-23761664-2384CONICET DigitalCONICETenginfo:eu-repo/semantics/altIdentifier/url/https://sjpp.springeropen.com/articles/10.1186/s13358-025-00402-4info:eu-repo/semantics/altIdentifier/doi/10.1186/s13358-025-00402-4info:eu-repo/semantics/openAccesshttps://creativecommons.org/licenses/by/2.5/ar/reponame:CONICET Digital (CONICET)instname:Consejo Nacional de Investigaciones Científicas y Técnicas2026-02-26T10:07:40Zoai:ri.conicet.gov.ar:11336/274803instacron:CONICETInstitucionalhttp://ri.conicet.gov.ar/Organismo científico-tecnológicoNo correspondehttp://ri.conicet.gov.ar/oai/requestdasensio@conicet.gov.ar; lcarlino@conicet.gov.arArgentinaNo correspondeNo correspondeNo correspondeopendoar:34982026-02-26 10:07:40.744CONICET Digital (CONICET) - Consejo Nacional de Investigaciones Científicas y Técnicasfalse
dc.title.none.fl_str_mv Testudinid turtle remains from the Late Miocene palaeo-island of Gargano, Italy, and an overview of Mediterranean insular tortoises
title Testudinid turtle remains from the Late Miocene palaeo-island of Gargano, Italy, and an overview of Mediterranean insular tortoises
spellingShingle Testudinid turtle remains from the Late Miocene palaeo-island of Gargano, Italy, and an overview of Mediterranean insular tortoises
Georgalis, Georgios L.
TURTLES
TESTUDINIDAE
ANATOMY
TAXONOMY
PALAEOGEOGRAPHY
INSULAR FAUNAS
title_short Testudinid turtle remains from the Late Miocene palaeo-island of Gargano, Italy, and an overview of Mediterranean insular tortoises
title_full Testudinid turtle remains from the Late Miocene palaeo-island of Gargano, Italy, and an overview of Mediterranean insular tortoises
title_fullStr Testudinid turtle remains from the Late Miocene palaeo-island of Gargano, Italy, and an overview of Mediterranean insular tortoises
title_full_unstemmed Testudinid turtle remains from the Late Miocene palaeo-island of Gargano, Italy, and an overview of Mediterranean insular tortoises
title_sort Testudinid turtle remains from the Late Miocene palaeo-island of Gargano, Italy, and an overview of Mediterranean insular tortoises
dc.creator.none.fl_str_mv Georgalis, Georgios L.
Vlachos, Evangelos
Camanni, Francesca
Villa, Andrea
Pavia, Marco
van den Hoek Ostende, Lars W.
Delfino, Massimo
author Georgalis, Georgios L.
author_facet Georgalis, Georgios L.
Vlachos, Evangelos
Camanni, Francesca
Villa, Andrea
Pavia, Marco
van den Hoek Ostende, Lars W.
Delfino, Massimo
author_role author
author2 Vlachos, Evangelos
Camanni, Francesca
Villa, Andrea
Pavia, Marco
van den Hoek Ostende, Lars W.
Delfino, Massimo
author2_role author
author
author
author
author
author
dc.subject.none.fl_str_mv TURTLES
TESTUDINIDAE
ANATOMY
TAXONOMY
PALAEOGEOGRAPHY
INSULAR FAUNAS
topic TURTLES
TESTUDINIDAE
ANATOMY
TAXONOMY
PALAEOGEOGRAPHY
INSULAR FAUNAS
purl_subject.fl_str_mv https://purl.org/becyt/ford/1.5
https://purl.org/becyt/ford/1
dc.description.none.fl_txt_mv The Late Miocene Gargano “Terre Rosse” in southern Italy has yielded a diverse vertebrate fauna that comprises an array of endemic taxa, including forms characterized by extreme sizes and unique morphologies. We here document new testudinid fossil remains from this insular assemblage. The material comprises a mandible, abundant (though mostly fragmentary) shell remains, appendicular elements, and a few vertebrae, originating from different localities within the Gargano assemblage. Based on its femur morphology, we tentatively refer the Gargano form to Solitudo, a late Neogene and Quaternary insular genus, otherwise known from a few Mediterranean islands. Unfortunately, the incomplete preservation of the trochanters hinders observation of the main diagnostic character of Solitudo, however, the similarities in the femoral head and diaphysis allow an open identification of the Gargano material as cf. Solitudo sp. Still, the Gargano tortoise offers new anatomical information on the Mediterranean insular tortoises. An interesting feature of the few preserved plastral elements is the presence of sulci as distinct raised ridges, a character that has been reported in other insular extant and extinct tortoises as well. The preserved hyoplastron has a medially straight and laterally concave humeropectoral sulcus and a wide axillary scute, pointing to some resemblance to geochelonans. Moreover, the morphology of the mandible from Gargano, which has its lingual serration reaching towards the symphyseal area, is also indicative of geochelonan affinities. If our identification is correct, the Gargano form would correspond to the earliest occurrence of Solitudo, denoting that this insular testudinid already radiated during the Late Miocene. A thorough overview of the diversity and taxonomy of insular testudinids from the Mediterranean islands is provided.
Fil: Georgalis, Georgios L.. Polish Academy Of Sciences. Institute Of Paleobiology; Argentina
Fil: Vlachos, Evangelos. Museo Paleontológico Egidio Feruglio; Argentina. Consejo Nacional de Investigaciones Científicas y Técnicas; Argentina
Fil: Camanni, Francesca. Università di Torino; Italia
Fil: Villa, Andrea. Institut Català de Paleontologia Miquel Crusafont.; España
Fil: Pavia, Marco. Università di Torino; Italia
Fil: van den Hoek Ostende, Lars W.. Naturalis Biodiversity Center; Países Bajos
Fil: Delfino, Massimo. Università di Torino; Italia
description The Late Miocene Gargano “Terre Rosse” in southern Italy has yielded a diverse vertebrate fauna that comprises an array of endemic taxa, including forms characterized by extreme sizes and unique morphologies. We here document new testudinid fossil remains from this insular assemblage. The material comprises a mandible, abundant (though mostly fragmentary) shell remains, appendicular elements, and a few vertebrae, originating from different localities within the Gargano assemblage. Based on its femur morphology, we tentatively refer the Gargano form to Solitudo, a late Neogene and Quaternary insular genus, otherwise known from a few Mediterranean islands. Unfortunately, the incomplete preservation of the trochanters hinders observation of the main diagnostic character of Solitudo, however, the similarities in the femoral head and diaphysis allow an open identification of the Gargano material as cf. Solitudo sp. Still, the Gargano tortoise offers new anatomical information on the Mediterranean insular tortoises. An interesting feature of the few preserved plastral elements is the presence of sulci as distinct raised ridges, a character that has been reported in other insular extant and extinct tortoises as well. The preserved hyoplastron has a medially straight and laterally concave humeropectoral sulcus and a wide axillary scute, pointing to some resemblance to geochelonans. Moreover, the morphology of the mandible from Gargano, which has its lingual serration reaching towards the symphyseal area, is also indicative of geochelonan affinities. If our identification is correct, the Gargano form would correspond to the earliest occurrence of Solitudo, denoting that this insular testudinid already radiated during the Late Miocene. A thorough overview of the diversity and taxonomy of insular testudinids from the Mediterranean islands is provided.
publishDate 2025
dc.date.none.fl_str_mv 2025-09
dc.type.none.fl_str_mv info:eu-repo/semantics/article
info:eu-repo/semantics/publishedVersion
http://purl.org/coar/resource_type/c_6501
info:ar-repo/semantics/articulo
format article
status_str publishedVersion
dc.identifier.none.fl_str_mv http://hdl.handle.net/11336/274803
Georgalis, Georgios L.; Vlachos, Evangelos; Camanni, Francesca; Villa, Andrea; Pavia, Marco; et al.; Testudinid turtle remains from the Late Miocene palaeo-island of Gargano, Italy, and an overview of Mediterranean insular tortoises; Springer; Swiss Journal of Palaeontology; 144; 1; 9-2025; 1-24
1664-2376
1664-2384
CONICET Digital
CONICET
url http://hdl.handle.net/11336/274803
identifier_str_mv Georgalis, Georgios L.; Vlachos, Evangelos; Camanni, Francesca; Villa, Andrea; Pavia, Marco; et al.; Testudinid turtle remains from the Late Miocene palaeo-island of Gargano, Italy, and an overview of Mediterranean insular tortoises; Springer; Swiss Journal of Palaeontology; 144; 1; 9-2025; 1-24
1664-2376
1664-2384
CONICET Digital
CONICET
dc.language.none.fl_str_mv eng
language eng
dc.relation.none.fl_str_mv info:eu-repo/semantics/altIdentifier/url/https://sjpp.springeropen.com/articles/10.1186/s13358-025-00402-4
info:eu-repo/semantics/altIdentifier/doi/10.1186/s13358-025-00402-4
dc.rights.none.fl_str_mv info:eu-repo/semantics/openAccess
https://creativecommons.org/licenses/by/2.5/ar/
eu_rights_str_mv openAccess
rights_invalid_str_mv https://creativecommons.org/licenses/by/2.5/ar/
dc.format.none.fl_str_mv application/pdf
application/pdf
dc.publisher.none.fl_str_mv Springer
publisher.none.fl_str_mv Springer
dc.source.none.fl_str_mv reponame:CONICET Digital (CONICET)
instname:Consejo Nacional de Investigaciones Científicas y Técnicas
reponame_str CONICET Digital (CONICET)
collection CONICET Digital (CONICET)
instname_str Consejo Nacional de Investigaciones Científicas y Técnicas
repository.name.fl_str_mv CONICET Digital (CONICET) - Consejo Nacional de Investigaciones Científicas y Técnicas
repository.mail.fl_str_mv dasensio@conicet.gov.ar; lcarlino@conicet.gov.ar
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score 13.176822

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